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Modifiers are applied to the melee strengths.
• If no attacking units have fired in that player’s turn, then 20% is added to the attacking strength.

We are playing PBEM with auto defensive fire. When does the above rule not trigger? Is there a case where a unit has fired and can attack in the same turn in the mode we play? You can only either attack in melee or fire afaik.

AFAIK the rule always applies. If you "waste" your fire before the melee you won't get that 20% bonus.

But that has nothing to do with what you can do, usually depending on the unit type, you can always fire & melee in the same turn.